They discovered that no regular cerevisiae or carlsbergensis strains utilised lysine whereas many different yeasts, together with wild yeasts, did so. They kept their stock cultures on malt extract agar slopes or on malt extract chalk agar within the case of Brettanomyces species. Later, Morris & Eddy1 described a strong lysine medium for the isolation and enumeration of wild yeasts in pitching yeast.

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