They found that no regular cerevisiae or carlsbergensis strains utilised lysine whereas many other yeasts, including wild yeasts, did so. They kept their stock cultures on malt extract agar slopes or on malt extract chalk agar in the case of Brettanomyces species. Later, Morris & Eddy1 described a strong lysine medium for the isolation and enumeration of untamed yeasts in pitching yeast.

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